| 2006-present: | Lecturer, University of Manchester. |
| 2005-2006: | Post-doctoral research fellow, EMBL - European Bioinformatics Institute. |
| 2002-2005: | Visiting scientist, EMBL - European Bioinformatics Institute. |
| 2000-2005: | Post-doctoral Research Associate, Department of Zoology, The University of Cambridge. |
| 1997-2000: | PhD on "Mathematical Modelling and Statistical Testing in Phylogenetics", supervised by Nick Goldman, Departments of Genetics and Zoology, The University of Cambridge |
| 1994-1997: | BSc(Hons) Genetics, The University of Nottingham |